Threats include disturbance from cavers and climbers, collection by bryologists, pollution of the spring in which it grows, and desiccation during periods of drought.
en.wikipedia.org The ability to clamp down also seals the shell edge against the rock surface, protecting them from desiccation during low tide, despite their being in full sunlight.
en.wikipedia.org The outer pollen wall, which prevents the pollen grain from shrinking and crushing the genetic material during desiccation, is composed of two layers.
en.wikipedia.org The breccia passes up into muddy sandstones, often with well-preserved desiccation structures, and into deposits of a braided river system, trough cross-bedded sandstones and conglomerates.
en.wikipedia.org This seals the opening of the shell and serves to protect the worm from predators and desiccation when out of water.
en.wikipedia.org