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The larger, older males establish and defend distinct territories on the rookery.
en.wikipedia.org While their rookeries had been thickly buried as well, coastal erosion had cleared them by the summer of 2009.
en.wikipedia.org Mating may occur outside the rookeries, between non-territorial males and females, as the latter move to and from the mating site.
en.wikipedia.org The area is hills that are mostly forested as well as a great blue heron rookery.
en.wikipedia.org Breeding takes place in these abandoned nests; usually breeding occurs colonially in rookeries because these birds tend to stay together in groups.
en.wikipedia.org